How to apply

We are part of the Coordinated Doctoral Programs in Biomedical Sciences at the Health Sciences Campus of Columbia University in the City of New York. These Programs emphasize research and training diverse areas of basic biomedical sciences. The distinguishing feature of each program is a distinct curriculum that is tailored to the overall programmatic aims.

The Coordinated Doctoral Programs are part of the Columbia University Graduate School of Arts and Sciences (GSAS) which includes graduate programs at both the Columbia University Irving Medical Center (CUIMC) and main Columbia campuses.

You must apply through the Biomedical Doctoral Programs.* Applications are generally open from mid-September to early December. 

Apply

Individual training programs are administered by separate committees and program directors. The program director for Genetics and Development are Mimi Shirasu-Hiza (ms4095@cumc.columbia.edu) and Luke Berchowitz (leb2210@cumc.columbia.edu). A training committee of five or six faculty oversees the training program. Current members, in addition to the program directors, are Laura Johnston, Chao Lu, and Joanna Smeeton. Please contact Jooyeon Kim (jk2857@cumc.columbia.edu) for training program administrative activities.

For more information, contact us

*Please be careful when applying. While we are a part of the Graduate School of Arts and Sciences, there are two applications. Make sure you are using the application for Columbia University Medical Center.